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Meristem parameters
The parameters in the Meristems section determine how meristems work, and therefore how the plant branches and develops.
Axillary meristem and leaf arrangement
Plants are often classified by how many leaves they have at each node -- one (alternate), two (opposite), or more than two (whorled). PlantStudio
simulates the first two cases, which cover most plants. Which of these arrangements
you choose determines how many axillary meristems are available for branching
and for producing inflorescences, because axillary meristems form in the angles
between leaf and stem (axils). So a plant with an opposite leaf arrangement
has twice as many axillary meristems as a plant with an alternate leaf
arrangement.
Meristem 3D object, scale, colors
These parameters determine how meristems appear on the plant. Normally you
will not want to see meristems, so you will keep the Meristems: Meristem 3D object scale at full size parameter set to zero. But if you want to see how the meristems work, or if
you are making some plant on which little buds would look right, you can
increase the scale.
Branching index, Apical dominance strength, Secondary branching
Branching in PlantStudio works in this way: Each day, each axillary meristem
that is not already committed to reproductive growth ponders whether it might
start a new branch. In making this decision the meristem takes three things into
account.
First, the meristem considers the branching index for the whole plant, which
runs from zero to one hundred. This gives the meristem a probabilistic limit to
check -- if it draws a random number between zero and one hundred and the number falls
below the branching index, the meristem can create a branch.
Second, the meristem may lower the branching index it uses based on its
proximity to an apical meristem. One of the ways plants control their shape is by apical dominance. In apical dominance the apical meristem at the end of the plant stem dominates the other meristems by sending out a substance that inhibits them from
branching. (This is why garden books tell you to "top" plants you want to grow branches -- you are removing the apical meristem.) PlantStudio simulates apical dominance
by reducing the probability of branching in meristems close to the stem apex.
You can control where the apical dominance effect starts by setting the
parameter Meristems: Apical dominance strength (as node distance). For example, if a plant stem has five nodes and you set the apical dominance
strength to 3.0, the first two nodes will branch normally but the last three
(closest to the apical meristem) will face an increasing difficulty in branching.
Finally, the meristem may be prevented from branching if you have turned off
secondary branching and it is not on the plant's main stem.
Usually you will want to keep the branching index at less than 30%, because
above that percentage there can be so much branching that the plant takes a very
long time to draw. You can also control the amount of branching by changing how
many axillary meristems are available to contemplate branching. You can reduce
the number of axillary meristems by choosing alternate leaf arrangement and by
increasing the minimum days needed to create new internodes and leaves.
Sympodial branching
Most plants have monopodial branching, in which apical meristems extend stems
and axillary meristems create new branches. In sympodial branching, each apical
meristem dies off after it has made one internode, and one of the axillary
meristems it created takes over the main stem. You can see this type of branching
in tomato plants. To specify sympodial branching, just turn this parameter on.
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| monopodial branching
| sympodial branching
| Branching angle
The branching angle parameter determines what angle new branches make with the
stems they developed out of. Because PlantStudio doesn't have a gravity model, setting this parameter to a large angle may mean that
you get branches at impossible angles. So it is best to keep the angle below 90
degrees unless you have little branching or disallow secondary branching.
Probability each meristem will become reproductive when gets flowering signal
This parameter is also called the determinate probability. When determinate
plants begin reproduction, they go all out and never grow vegetatively again.
Indeterminate plants, on the other hand, still grow vegetatively though most of
their resources go to reproduction. You may have seen this distinction among
tomato varieties: bush varieties tend to be determinate (they grow, then flower,
then die), while vine varieties are usually indeterminate (they grow, then flower
and grow, then flower and grow some more, and they may be kept alive and
growing for a long time in a controlled environment). PlantStudio simulates that
difference here using a determinate probability, which is the probability that any
one meristem will switch over to reproduction when the plant enters its
reproductive phase. A determinate probability of one means that the plant makes no
more vegetative parts. A determinate probability of zero means the plant will
never flower. A determinate probability of 0.5 means that about half the meristems
will switch over and half will keep growing vegetatively.
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